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Centre for Development, Environment and Policy (CeDEP)

C191 Assessing and Understanding Biological Diversity

Course Code:
C191
Unit value:

Less than a seventh of the estimated number of species alive today have been scientifically described and, of these, the interrelationships between them are only understood for a fraction. In other words, there is a dearth of knowledge about biological diversity (biodiversity) and how individual taxa are related to each other. Detailed knowledge about populations, their habitats, and how species interact within ecosystems is also in short supply – a vast amount of work is still to be undertaken by taxonomists and conservation scientists before careless human action results in further biodiversity loss.

Biodiversity conservation and sustainable use is dependent on knowledge of the ecological functioning, biology, distribution, genetics, and taxonomy of organisms. In addition, knowledge of ecosystem functions and processes, species richness, centres of diversity, levels of habitat loss and threats, are necessary to decide on priority areas, habitats and taxa for conservation. The need for this knowledge is enshrined in Article 7 of the Convention on Biological Diversity (CBD), which requires parties to ‘identify and monitor the components of biodiversity important for conservation and sustainable use, and the processes and activities that are having or are likely to have a significant adverse impact on the conservation and sustainable use of biodiversity’. In addition, Parties are required to ‘maintain and organise the data derived from identification and monitoring activities’. To gain this knowledge and satisfy this requirement of the CBD, we must find ways of measuring, assessing and classifying all levels of biodiversity: genes, species and ecosystems.

This module presents the backbones of genetic conservation, approaches to inventorying and monitoring biodiversity and applied taxonomy. It explains the general principles of genetic diversity, outlining the theory of population genetics, and methods of measuring genetic, population, species, and ecosystem diversity are discussed. An overview of the variety of life forms is provided and the main elements of taxonomy are outlined. The major types of classification and means of identifying biodiversity are reviewed.

Objectives and learning outcomes of the course

Module Aims

Assessing and understanding biodiversity is a subject area that draws on a wide range of underlying theory and practice. It is therefore multi-disciplinary in nature and this module is essentially an introduction that maps out the different dimensions of the subject. While each unit is self-contained, the module is generally designed so that knowledge gained from one unit is used when studying subsequent units. Where prior knowledge is specifically needed before studying a unit, a note is included in the unit overview. For example, before studying Unit 4, Genetic conservation, you should be familiar with techniques for measuring genetic diversity (Unit 3) and complementary conservation strategies (in situ and ex situ conservation), which is covered in module C117, Introduction to Biodiversity Conservation.

The module aims to provide the minimum knowledge required for individuals planning a career in biodiversity conservation or a related field, or for those already involved in this area. The information presented in this module is equally relevant to policy makers as it is to field scientists because it is important that those in decision making positions are equipped with an understanding of the complexity of biodiversity assessment and of how demanding and resource-intensive it is. The module is also relevant to those with an interest in biodiversity conservation as a hobby rather than on a professional basis.

The specific aims of the module are to:

  • provide a background in the principles of genetic conservation
  • convey methods of assessing and measuring biodiversity at genetic, population, species and ecosystem levels
  • review the variety of life forms
  • provide a background in applied taxonomy and an understanding of its importance in biodiversity conservation
Module Learning Outcomes
  • explain how the fundamental principles of genetic conservation are applied in biodiversity conservation planning and implementation
  • identify a range of different methods available for measuring genetic, species and ecosystem diversity and make informed choices about which techniques would be appropriate in specific circumstances
  • explain to the person sitting next to you (whether at work, at home or on the bus) the vast variety of life forms that are known to exist on our planet and why taxonomy is fundamental to the conservation and management of this diversity
  • plan a taxonomic collecting expedition
  • explain why identification is critical for effective biodiversity conservation and find and use appropriate identification aids
  • make use of existing taxonomic data sources in conservation planning and outline how and why taxonomic revisions are undertaken.

Scope and syllabus

The module is arranged in four parts:

Part I

Part I provides a general introduction and overview of the assessment of biodiversity at each of three levels: genes, species and ecosystems.

Part II

Part II addresses the assessment of biodiversity at the genetic level. For those with little or no background in genetics, the first unit in this part provides an introduction to the principles of genetics. The second unit explains the tools and techniques available to measure genetic diversity between individuals and taxa, and within and between populations. The unit focuses on the use of molecular markers for studying diversity, outlining the different techniques available. The third unit reviews the role that genetic diversity studies play in biodiversity conservation and describes genetic conservation – the study of genetic variation within and between populations and taxa and its subsequent use in the formulation and implementation of conservation strategies.

Part III

Part III tackles the subject of inventorying and monitoring, which provides fundamental and essential biological information for biodiversity conservation, as reflected in Article 7 of the CBD. Inventorying and monitoring can be carried out at different spatial scales and at different biological levels. Careful planning of inventorying and monitoring programmes is crucial to the success of any project – clearly defined goals must be made before designing the monitoring strategy. Many methods can be used to inventory and monitor biodiversity and it is important to choose the appropriate method in order to answer the questions being posed. These issues are addressed in this part, which is divided logically into two units: the first focuses on inventory and monitoring of populations and species, while the second addresses ecosystems.

Part IV

Part IV is devoted to taxonomy – the study and description of variation patterns between organisms and the manipulation of the data to produce a system of classification. Taxonomy, through the organism naming system it provides, contributes the referencing system for the whole of biology and provides the backbone on to which pieces of biological information, including conservation data, are attached and communicated. An understanding of taxonomy is fundamental to the formulation and implementation of effective biodiversity conservation programmes. This part comprises four units: the first introduces the subject of taxonomy and provides a brief overview of the variety of life forms that exist on our planet. The second unit explains how and why taxonomic research is undertaken, with a focus on the process and products of taxonomic revisions – the means by which new taxa, classifications and associated information are produced and published. The third unit provides a broad review of two critical aspects of biodiversity conservation: taxonomic collection and identification – arguably the most important facets of taxonomy for conservationists. The unit discusses the principles of collecting taxonomic reference samples and reviews the various forms of identification aid. The fourth and final unit explains how taxonomic data is analysed and provides an overview of sources of taxonomic information.

Suggested reading

Study guide

Your study guide, on CD, comprises:

  • 10 module units
  • 2 Tutor Marked Assignments (TMAs)
  • self-assessment questions and answers
  • in-text questions and activities
  • a specimen exam paper

We strongly advise that you read the sections on study methods and using the module CD prior to study.

Textbooks

Two textbooks have been selected to accompany your study guide. Please note that you are not expected to read these texts in their entirety. They are provided to elaborate and build on the contents of the study guide and thus to deepen your understanding of the subject. You may also find them useful for reference purposes; for example, to look up a particular subject that you would like to read about in more depth or to look up the meaning of specific terms. In many cases, the unit key readings (see below) are drawn from the accompanying texts.

  • Allendorf FW and Luikart G (2006) Conservation and the Genetics of Populations, Blackwell Publishing, ISBN 1405121459, Softcover
    This text provides a comprehensive overview of the essential background, concepts, and tools needed to understand how genetic information can be used to develop species conservation plans. Using a balance of data and theory, and basic and applied research, the book examines genetic and phenotypic variation in natural populations, the principles and mechanisms of evolutionary change, the interpretation of genetic data from natural populations and how this may be applied to conservation. The text includes a very useful glossary of terms and definitions of mathematical symbols.
    In addition to providing general reading material to accompany your study guide, this text also provides some of the key readings for Units 1-4 of the module. You will notice that this text includes a lot of mathematical equations and graphs. You should not be concerned if you do not fully understand these. From reading this text and other readings that accompany the module, it is important to absorb key concepts, not mathematical equations or minute details. Try to stay focused on the broad concepts being presented as you read; these are what is important. However, if you are planning to undertake further research in a particular field of genetic conservation, you will then find a deeper understanding of the underlying mathematics useful.
  • Tudge C (2000) The Variety of Life: A Survey and Celebration of all the Creatures that have ever Lived, Oxford University Press, ISBN 0198604262, Softcover
    Tudge provides the most comprehensive text available on the variety of life. This text provides a summary of all creatures, alive and extinct, and shows the principal groups to which all living things belong. Whilst you are not expected to have an in-depth knowledge of all the biological groups, it is important to comprehend the breadth of biodiversity in order to appreciate the enormity of the task of assessing, understanding, conserving, and managing it. The text provides a very good overview of the history and development of biological classification, presenting key taxonomic concepts and methods, as well as reviewing the types of data that can be used, which provides reading material to support units 7–10 of the module.
    Key readings
    Each unit comprises a set of key readings from books, journals and other publications; these form an integral part of the module and provide a deeper understanding of the subjects covered in the study guide. You will be expected to draw on knowledge gained from the key readings in the exams; however, you will not be examined directly on the content of these readings.
    The section on the CD listing the key readings gives brief overviews of the readings and how they complement the unit material. The readings are provided in bound volumes, and if a reading relates to more than one unit, you will find that a copy is provided only for the first unit that the reading appears in.
Further readings and references

Each unit also contains a list of further resources and references which will help your understanding and appreciation of the complexities of the subject matter. These readings are not integral to the module and are therefore not provided in your study pack; they are optional.

Additional resources

In addition to references relating to specific units, below is a list of selected resources that you may find helpful in the overall context of the module. This list is not, however, exhaustive.

Textbooks

A full list of textbooks can be found in the downloadable pdf: C191: Full Details